What causes 'astringency' — a harsh, drying, puckering sensation — in finished beer?
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A
Excessive polyphenol/tannin extraction from over-sparging or grain husks at high sparge temperatures
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B
Excessive hop rates beyond the style's appropriate IBU range
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C
High carbonation levels irritating the palate
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D
Flocculent yeast strains leaving protein haze in the beer