A student is revising the following passage about ecological succession:
"Primary succession begins on barren substrates devoid of soil. Lichens colonize bare rock. They secrete acids that slowly break down rock into mineral particles. Over centuries, these particles accumulate with organic matter from dead lichens. Soil eventually forms."
Which version most effectively combines the underlined sentences into a single, structurally sophisticated sentence?
"Lichens colonize bare rock. They secrete acids that slowly break down rock into mineral particles."
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A
Lichens colonize bare rock, and they secrete acids that slowly break down rock into mineral particles.
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B
Lichens, colonizing bare rock, secrete acids that slowly break down rock into mineral particles.
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C
Colonizing bare rock, lichens' acid secretions slowly break down the rock into mineral particles.
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D
Bare rock is colonized by lichens, which slowly break it down into mineral particles through acid secretions.